A cross is made between two E. coli strains: Hfr arg⁺ bio⁺ leu⁺ × F arg⁻ bio⁻ leu⁻. Inter⁻ rupted mating studies show that argt enters the recipient last, and so arg⁺ recombi- nants are selected on a medium containing bio and leu only. These recombinants are tested for the presence of bio⁺ and leu⁺. The following numbers of individuals are found for each genotype:

arg⁺ bio⁺ leu⁺ 320 arg bio leu+ 0
arg⁺ bio⁺ leu⁻ 8 arg⁺ bio⁻ leu⁻ 48

What are the map distances in recombination percentages?